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Evolutions

Evolving 3 items in one chest.

Evolution is a game mechanic that typically changes a base weapon to a stronger evolved weapon using a passive item as a catalyst. A Union is a combination of two weapons. A Gift is an additional weapon or passive item you would get when the conditions were met.

Evolved weapons usually inherit most of the base weapon stats with extra effects, most Area-of-Effect weapons may add bonus to Area and some have changes on base damage because their behavior is different. United weapons share similar characteristics with evolved weapons, except two weapons are merged into one, hence emptying a weapon slot after the united weapon is obtained. Gifts act as an additional weapon or passive item, without replacing the base weapon.

Mechanics[]

To evolve or unite a weapon, or to obtain a gift, the player needs to fulfill following conditions:

  • The player must have a fully leveled base weapon and its counterpart (required level varies by case) in their inventory.
  • The player must then collect a Treasure Chest that contains an evolution type reward. In most stages, this kind of chests are dropped by bosses that spawned at or after the 10-minute mark with some exceptions:
    • The first chest in Mad Forest dropped by the Glowing Bat, which spawns along the wave at 1:00, can evolve a weapon.
    • All chests in Dairy Plant, Il Molise, Cappella Magna, Boss Rash, Laborratory, Abyss Foscari Tides Of The Foscari DLC, and Hectic Highway Operation Guns DLC can evolve a weapon regardless of time.
    • In Green Acres the ability to evolve a weapon is the same as the wave in its original stage.
      • For example, chests dropped by waves from Dairy Plant prior to 10 minute mark can evolve a weapon, but waves from Inlaid Library cannot.
    • In Tiny Bridge, all chests, except the chests from minute 3:00 and 6:00 bosses, can evolve a weapon.
    • At Mt.Moonspell Legacy Of The Moonspell DLC, all chests, except the chests from the minute 3:00 and 5:00 bosses, can evolve a weapon.
    • The first chest in Lake Foscari Tides Of The Foscari DLC dropped by the Foscaritrice, which spawns at 2:00, can evolve a weapon.
    • In Neo Galuga Operation Guns DLC, the chest dropped by Metal Alien at 9:00 can evolve a weapon.
    • Treasure Chests that contain an Arcana (minutes 11:00 and 21:00 in every normal stage) can only evolve a weapon when the player is already at the max count of Arcana.
    • The Character Gyorunton can evolve weapons with any chest pickup that gives an item, regardless of other conditions.

The chest will then grant an evolved weapon as its first reward in place of the usual random upgrade. On evolution, the base weapon is typically removed and the evolved weapon is added. The passive item remains unaffected. On union, both base weapons are removed and their union is added, freeing a weapon slot. On gift, the base weapon will not be replaced when gift is obtained.

Generally, only one weapon can be evolved per chest, including the higher tier chests. However, some chests contain multiple evolutions:

Chests in Il Molise, Bat Country, and Tiny Bridge, and the chest from Giant Enemy Crab can evolve up to five weapons. The rest can evolve up to three. To evolve multiple weapons at once, these chests must be of the higher tier with enough rewards, as evolution is a type of reward.

When playing in Co-op, weapons are capable of evolution regardless of which player has the Passive Weapon and which player has the Active Weapon, so long as the requirements are met and the share passives option in the level select menu is enabled.

Morph[]

Certain weapons evolve automatically when a character morphs into a new form, which occurs if you have the required relic and meet the required conditions to morph. When a character morphs, their starting weapon evolves into a new weapon as well. Picking up a treasure chest is not required.

Currently, only Mortaccio, Yatta Cavallo, and O'Sole Meeo can morph.

List of evolutions, unions, and gifts[]

Evolutions[]

Base weapon Passive item Evolution
Sprite-Whip
Whip
+ Sprite-Hollow Heart
Hollow Heart
= Sprite-Bloody Tear
Bloody Tear
Sprite-Magic Wand
Magic Wand
+ Sprite-Empty Tome
Empty Tome
= Sprite-Holy Wand
Holy Wand
Sprite-Knife
Knife
+ Sprite-Bracer
Bracer
= Sprite-Thousand Edge
Thousand Edge
Sprite-Axe
Axe
+ Sprite-Candelabrador
Candelabrador
= Sprite-Death Spiral
Death Spiral
Sprite-Cross
Cross
+ Sprite-Clover
Clover
= Sprite-Heaven Sword
Heaven Sword
Sprite-King Bible
King Bible
+ Sprite-Spellbinder
Spellbinder
= Sprite-Unholy Vespers
Unholy Vespers
Sprite-Fire Wand
Fire Wand
+ Sprite-Spinach
Spinach
= Sprite-Hellfire
Hellfire
Sprite-Garlic
Garlic
+ Sprite-Pummarola
Pummarola
= Sprite-Soul Eater
Soul Eater
Sprite-Santa Water
Santa Water
+ Sprite-Attractorb
Attractorb
= Sprite-La Borra
La Borra
Sprite-Runetracer
Runetracer
+ Armor
Armor
= Sprite-NO FUTURE
NO FUTURE
Sprite-Lightning Ring
Lightning Ring
+ Sprite-Duplicator
Duplicator
= Sprite-Thunder Loop
Thunder Loop
Sprite-Pentagram
Pentagram
+ Sprite-Crown
Crown
= Sprite-Gorgeous Moon
Gorgeous Moon
Sprite-Gatti Amari
Gatti Amari
+ Sprite-Stone Mask
Stone Mask
= Sprite-Vicious Hunger
Vicious Hunger
Sprite-Song of Mana
Song of Mana
+ Sprite-Skull O'Maniac
Skull O'Maniac
= Sprite-Mannajja
Mannajja
Sprite-Shadow Pinion
Shadow Pinion
+ Sprite-Wings
Wings
= Sprite-Valkyrie Turner
Valkyrie Turner
Sprite-Flames of Misspell
Flames of Misspell
+ Max level:
Sprite-Torrona's Box
Torrona's Box
= Sprite-Ashes of Muspell
Ashes of Muspell
Sprite-Glass Fandango
Glass Fandango
+ Max level:
Sprite-Wings
Wings
= Sprite-Celestial Voulge
Celestial Voulge
Sprite-Clock Lancet
Clock Lancet
+ Max level:
Sprite-Silver Ring
Silver Ring
Sprite-Gold Ring
Gold Ring
= Sprite-Infinite Corridor
Infinite Corridor
Sprite-Laurel
Laurel
+ Max level:
Sprite-Metaglio Left
Metaglio Left
Sprite-Metaglio Right
Metaglio Right
= Sprite-Crimson Shroud
Crimson Shroud
Sprite-Phas3r
Phas3r
+ Max level:

Sprite-Empty Tome
Empty Tome

= Sprite-Photonstorm
Photonstorm
Sprite-Pako Battiliar
Pako Battiliar
+ Max level:

Sprite-Hollow Heart
Hollow Heart

= Sprite-Mazo Familiar
Mazo Familiar
Sprite-Santa Javelin
Santa Javelin
+ Max level:

Sprite-Clover
Clover

= Sprite-Seraphic Cry
Seraphic Cry
Base weapon Evolution
Sprite-Bracelet
Bracelet
= Sprite-Bi-Bracelet
Bi-Bracelet
Sprite-Bi-Bracelet
Bi-Bracelet
= Sprite-Tri-Bracelet
Tri-Bracelet

Unions[]

Base weapon Passive item Evolution & Union
Sprite-Phiera Der Tuphello
Phiera Der Tuphello
Sprite-Eight The Sparrow
Eight The Sparrow
+ Sprite-Tirajisú
Tirajisú
= Sprite-Phieraggi
Phieraggi
Base weapon Base weapon Union
Sprite-Peachone
Peachone
+ Sprite-Ebony Wings
Ebony Wings
= Sprite-Vandalier
Vandalier
Sprite-Vento Sacro
Vento Sacro
+ Sprite-Bloody Tear
Bloody Tear
= Sprite-Fuwalafuwaloo
Fuwalafuwaloo

Gifts[]

Base weapon Passive item Gift
Sprite-Victory Sword
Victory Sword
+ Max level:
Sprite-Torrona's Box
Torrona's Box
= Sprite-Sole Solution
Sole Solution
Base weapon Gift
Sprite-Candybox
Candybox
= Sprite-Super Candybox II Turbo
Super Candybox II Turbo

Morph[]

Base weapon Character Relic Condition(s) Evolution
Sprite-Bone
Bone
+ Sprite-Mortaccio
Mortaccio
+ Sprite-Chaos Malachite
Chaos Malachite
+ Reach level 80 = Sprite-Anima of Mortaccio
Anima of Mortaccio
Sprite-Cherry Bomb
Cherry Bomb
+ Sprite-Cavallo
Cavallo
+ Sprite-Chaos Rosalia
Chaos Rosalia
+ Reach level 80 = Sprite-Yatta Daikarin
Yatta Daikarin
Sprite-Celestial Dusting
Celestial Dusting
+ Sprite-O'Sole
O'Sole
+ Sprite-Chaos Altemanna
Chaos Altemanna
+ Reach level 80 = Sprite-Profusione D'Amore
Profusione D'Amore

DLC[]

Legacy of the Moonspell[]

Base weapon Passive item Evolution
Sprite-Silver Wind
Silver Wind Legacy Of The Moonspell DLC
+ Max level:
Sprite-Pummarola
Pummarola
= Sprite-Festive Winds
Festive Winds Legacy Of The Moonspell DLC
Sprite-Four Seasons
Four Seasons Legacy Of The Moonspell DLC
+ Max level:
Sprite-Candelabrador
Candelabrador
= Sprite-Godai Shuffle
Godai Shuffle Legacy Of The Moonspell DLC
Sprite-Summon Night
Summon Night Legacy Of The Moonspell DLC
+ Max level:
Sprite-Duplicator
Duplicator
= Sprite-Echo Night
Echo Night Legacy Of The Moonspell DLC
Sprite-Mirage Robe
Mirage Robe Legacy Of The Moonspell DLC
+ Max level:
Sprite-Attractorb
Attractorb
= Sprite-J'Odore
J'Odore Legacy Of The Moonspell DLC
Sprite-Night Sword
Night Sword Legacy Of The Moonspell DLC
+ Max level:
Sprite-Stone Mask
Stone Mask
= Sprite-Muramasa
Muramasa Legacy Of The Moonspell DLC
Sprite-Mille Bolle Blu
Mille Bolle Blu Legacy Of The Moonspell DLC
+ Max level:
Sprite-Spellbinder
Spellbinder
= Sprite-Boo Roo Boolle
Boo Roo Boolle Legacy Of The Moonspell DLC

Tides of the Foscari[]

Base weapon Passive item Evolution/Union
Sprite-SpellString
SpellString Tides Of The Foscari DLC
Sprite-SpellStream
SpellStream Tides Of The Foscari DLC
Sprite-SpellStrike
SpellStrike Tides Of The Foscari DLC
= Sprite-SpellStrom
SpellStrom Tides Of The Foscari DLC
Sprite-Eskizzibur
Eskizzibur Tides Of The Foscari DLC
+ Max level:
Sprite-Armor
Armor
= Sprite-Legionnaire
Legionnaire Tides Of The Foscari DLC
Sprite-Flash Arrow
Flash Arrow Tides Of The Foscari DLC
+ Max level:
Sprite-Bracer
Bracer
= Sprite-Millionaire
Millionaire Tides Of The Foscari DLC
Sprite-Prismatic Missile
Prismatic Missile Tides Of The Foscari DLC
+ Max level:
Sprite-Crown
Crown
= Sprite-Luminaire
Luminaire Tides Of The Foscari DLC
Sprite-Shadow Servant
Shadow Servant Tides Of The Foscari DLC
+ Max level:
Sprite-Skull O'Maniac
Skull O'Maniac
= Sprite-Ophion
Ophion Tides Of The Foscari DLC

Emergency Meeting[]

The passive item will be consumed while evolving, and can be obtained again though leveling, or pick-ups.

Base weapon Passive item Evolution/Union
Sprite-Report!
Report! Emergency Meeting DLC
+ Max level:
Sprite-Mini Crewmate
Mini Crewmate Emergency Meeting DLC
= Emergency Meeting
Emergency Meeting Emergency Meeting DLC
Sprite-Lucky Swipe
Lucky Swipe Emergency Meeting DLC
+ Max level:
Sprite-Mini Engineer
Mini Engineer Emergency Meeting DLC
= Sprite-Crossed Wires
Crossed Wires Emergency Meeting DLC
Sprite-Lifesign Scan
Lifesign Scan Emergency Meeting DLC
+ Max level:
Sprite-Mini Ghost
Mini Ghost Emergency Meeting DLC
= Sprite-Paranormal Scan
Paranormal Scan Emergency Meeting DLC
Sprite-Just Vent
Just Vent Emergency Meeting DLC
+ Max level:
Sprite-Mini Shapeshifter
Mini Shapeshifter Emergency Meeting DLC
= Sprite-Unjust Ejection
Unjust Ejection Emergency Meeting DLC
Sprite-Clear Debris
Clear Debris Emergency Meeting DLC
+ Max level:
Sprite-Mini Guardian
Mini Guardian Emergency Meeting DLC
= Sprite-Clear Asteroids
Clear Asteroids Emergency Meeting DLC
Sprite-Sharp Tongue
Sharp Tongue Emergency Meeting DLC
+ Max level:
Sprite-Mini Impostor
Mini Impostor Emergency Meeting DLC
= Sprite-Impostongue
Impostongue Emergency Meeting DLC
Sprite-Science Rocks
Science Rocks Emergency Meeting DLC
+ Max level:
Sprite-Mini Scientist
Mini Scientist Emergency Meeting DLC
= Sprite-Rocket Science
Rocket Science Emergency Meeting DLC


Operation Guns[]

Base weapon Passive item Passive item Evolution/Union
Sprite-Long Gun
Long Gun Operation Guns DLC
+ Sprite-Weapon Power-Up
Weapon Power-Up Operation Guns DLC
= Sprite-Prototype A
Prototype A Operation Guns DLC
Sprite-Short Gun
Short Gun Operation Guns DLC
+ Sprite-Weapon Power-Up
Weapon Power-Up Operation Guns DLC
+ Sprite-Bracer
Bracer
= Sprite-Prototype B
Prototype B Operation Guns DLC
Sprite-Spread Shot
Spread Shot Operation Guns DLC
+ Sprite-Weapon Power-Up
Weapon Power-Up Operation Guns DLC
+ Sprite-Empty Tome
Empty Tome
= Sprite-Prototype C
Prototype C Operation Guns DLC
Sprite-C-U-Laser
C-U-Laser Operation Guns DLC
+ Sprite-Weapon Power-Up
Weapon Power-Up Operation Guns DLC
+ Sprite-Tirajisú
Tirajisú
= Sprite-Pronto Beam
Pronto Beam Operation Guns DLC
Sprite-Firearm
Firearm Operation Guns DLC
+ Sprite-Weapon Power-Up
Weapon Power-Up Operation Guns DLC
+ Sprite-Candelabrador
Candelabrador
= Sprite-Fire-L3GS
Fire-L3GS Operation Guns DLC
Sprite-Sonic Bloom
Sonic Bloom Operation Guns DLC
+ Sprite-Weapon Power-Up
Weapon Power-Up Operation Guns DLC
+ Sprite-Armor
Armor
= Sprite-Wave Beam
Wave Beam Operation Guns DLC
Sprite-Homing Miss
Homing Miss Operation Guns DLC
+ Sprite-Weapon Power-Up
Weapon Power-Up Operation Guns DLC
+ Sprite-Duplicator
Duplicator
= Sprite-Multistage Missiles
Multistage Missiles Operation Guns DLC
Sprite-Diver Mines
Diver Mines Operation Guns DLC
+ Sprite-Weapon Power-Up
Weapon Power-Up Operation Guns DLC
+ Sprite-Attractorb
Attractorb
= Sprite-Atmo-Torpedo
Atmo-Torpedo Operation Guns DLC
Sprite-Blade Crossbow
Blade Crossbow Operation Guns DLC
+ Sprite-Weapon Power-Up
Weapon Power-Up Operation Guns DLC
+ Sprite-Clover
Clover
= Sprite-BFC2000-AD
BFC2000-AD Operation Guns DLC
Sprite-Prism Lass
Prism Lass Operation Guns DLC
+ Sprite-Weapon Power-Up
Weapon Power-Up Operation Guns DLC
+ Sprite-Wings
Wings
= Sprite-Time Warp
Time Warp Operation Guns DLC
Sprite-Metal Claw
Metal Claw Operation Guns DLC
+ Sprite-Weapon Power-Up
Weapon Power-Up Operation Guns DLC
+ Sprite-Hollow Heart
Hollow Heart
= Sprite-Big Fuzzy Fist
Big Fuzzy Fist Operation Guns DLC


Ode to Castlevania[]

Base weapon Passive item Evolution
Sprite-Alchemy Whip
Alchemy Whip
+ Sprite-Tirajisú
Tirajisú
= Sprite-Vampire Killer
Vampire Killer
Sprite-Wind Whip
Wind Whip
+ Sprite-Crown
Crown
= Sprite-Spirit Tornado Tip
Spirit Tornado Tip
Sprite-Platinum Whip
Platinum Whip
+ Sprite-Clover
Clover
= Sprite-Cross Crasher Tip
Cross Crasher Tip
Sprite-Dragon Water Whip
Dragon Water Whip
+ Sprite-Attractorb
Attractorb
= Sprite-Hydrostormer Tip
Hydrostormer Tip
Sprite-Sonic Whip
Sonic Whip
+ Sprite-Skull O'Maniac
Skull O'Maniac
= Sprite-Crissaegrim Tip
Crissaegrim Tip
Sprite-Jet Black Whip
Jet Black Whip
+ Sprite-Stone Mask
Stone Mask
= Sprite-Mormegil Tip
Mormegil Tip
Sprite-Vibhuti Whip
Vibhuti Whip
+ Sprite-Candelabrador
Candelabrador
= Sprite-Daybreaker Tip
Daybreaker Tip
Sprite-Vanitas Whip
Vanitas Whip
+ Sprite-Hollow Heart
Hollow Heart
= Sprite-Aurablaster Tip
Aurablaster Tip
Sprite-Shuriken
Shuriken
+ Sprite-Empty Tome
Empty Tome
= Sprite-Yagyu Shuriken
Yagyu Shuriken
Sprite-Curved Knife
Curved Knife
+ Sprite-Bracer
Bracer
= Sprite-Bwaka Knife
Bwaka Knife
Sprite-Javelin
Javelin
+ Sprite-Spellbinder
Spellbinder
= Sprite-Long Inus
Long Inus
Sprite-Discus
Discus
+ Max level:

Sprite-Parm Aegis
Parm Aegis

= Sprite-Stellar Blade
Stellar Blade
Sprite-Iron Ball
Iron Ball
+ Sprite-Armor
Armor
= Sprite-Wrecking Ball
Wrecking Ball
Sprite-Silver Revolver
Silver Revolver
+ Sprite-Karoma's Mana
Karoma's Mana
= Sprite-Jewel Gun
Jewel Gun
Sprite-Hand Grenade
Hand Grenade
+ Max level:

Sprite-Candelabrador
Candelabrador

= Sprite-The RPG
The RPG
Sprite-Wine Glass
Wine Glass
+ Max level:

Sprite-Tirajisú
Tirajisú

= Sprite-Meal Ticket
Meal Ticket
Sprite-Raging Fire
Raging Fire
+ Max level:

Sprite-Spinach
Spinach

= Sprite-Salamender
Salamender
Sprite-Ice Fang
Ice Fang
+ Max level:
Sprite-Spellbinder
Spellbinder
= Sprite-Cocytus
Cocytus
Sprite-Gale Force
Gale Force
+ Max level:

Sprite-Bracer
Bracer

= Sprite-Pneuma Tempestas
Pneuma Tempestas
Sprite-Rock Riot
Rock Riot
+ Max level:
Sprite-Stone Mask
Stone Mask
= Sprite-Gemma Torpor
Gemma Torpor
Sprite-Fulgur
Fulgur
+ Max level:

Sprite-Duplicator
Duplicator

= Sprite-Tenebris Tonitrus
Tenebris Tonitrus
Sprite-Keremet Bubbles
Keremet Bubbles
+ Max level:

Sprite-Armor
Armor

= Sprite-Keremet Morbus
Keremet Morbus
Sprite-Hex
Hex
+ Max level:

Sprite-Skull O'Maniac
Skull O'Maniac

= Sprite-Nightmare
Nightmare
Sprite-Refectio
Refectio
+ Max level:

Sprite-Clover
Clover

= Sprite-Sanctuary
Sanctuary
Sprite-Mace
Mace
+ Max level:

Sprite-Hollow Heart
Hollow Heart

= Sprite-Stamazza
Stamazza
Sprite-Star Flail
Star Flail
+ Max level:

Sprite-Pummarola
Pummarola

= Sprite-Moon Rod
Moon Rod
Sprite-Alucard Spear
Alucard Spear
+ Sprite-Wings
Wings
= Sprite-Thunderbolt Spear
Thunderbolt Spear
Sprite-Trident
Trident
+ Max level:

Sprite-Duplicator
Duplicator

= Sprite-Gungnir-Souris
Gungnir-Souris
Sprite-Iron Shield
Iron Shield
+

Sprite-Parm Aegis
Parm Aegis

= Sprite-Dark Iron Shield
Dark Iron Shield
Sprite-Guardian's Targe
Guardian's Targe
+ Sprite-Pummarola
Pummarola
= Sprite-Sacred Beasts Tower Shield
Sacred Beasts Tower Shield
Sprite-Tyrfing
Tyrfing
+ Sprite-Spinach
Spinach
= Sprite-Rune Sword
Rune Sword
Sprite-Alucart Sworb
Alucart Sworb
= Sprite-Alucard Swords
Alucard Swords
Sprite-Alucard Swords
Alucard Swords
+ 6
passives
&
evolutions
= Sprite-Alucard Shield
Alucard Shield
Sprite-Confodere
Confodere
= Sprite-Vol Confodere
Vol Confodere
Sprite-Vol Confodere
Vol Confodere
= Sprite-Melio Confodere
Melio Confodere
Sprite-Globus
Globus
+ Max level:

Sprite-Empty Tome
Empty Tome

= Sprite-Nitesco
Nitesco
Sprite-Optical Shot
Optical Shot
+ Max level:

Sprite-Karoma's Mana
Karoma's Mana

= Sprite-Acerbatus
Acerbatus
Sprite-Centralis Custos
Centralis Custos
Sprite-Dextro Custos
Dextro Custos
Sprite-Sinestro Custos
Sinestro Custos
= Sprite-Trinum Custodem
Trinum Custodem
Sprite-Dominus Agony
Dominus Agony
Sprite-Dominus Anger
Dominus Anger
Sprite-Dominus Hatred
Dominus Hatred
= Sprite-Power of Sire
Power of Sire
Sprite-Sonic Dash
Sonic Dash
+ Max level:

Sprite-Wings
Wings

= Sprite-Rapidus Fio
Rapidus Fio
Sprite-Luminatio
Luminatio
+ Max level:

Sprite-Crown
Crown

= Sprite-Vol Luminatio
Vol Luminatio
Sprite-Umbra
Umbra
+ Max level:

Sprite-Attractorb
Attractorb

= Sprite-Vol Umbra
Vol Umbra
Sprite-Vol Luminatio
Vol Luminatio
Sprite-Vol Umbra
Vol Umbra
= Sprite-Universitas
Universitas
Sprite-Endo Gears
Endo Gears
Sprite-Peri Pendulum
Peri Pendulum
Sprite-Myo Lift
Myo Lift
Sprite-Epi Head
Epi Head
= Sprite-Clock Tower
Clock Tower


Trivia[]

  • With the addition of Vicious Hunger, Shadow Pinion and Valkyrie Turner in patch 0.5.2, all passive items at the time have an evolution.
  • With the addition of Silver Ring, Gold Ring, Metaglio Left, and Metaglio Right in patch 0.6.1, all base weapons, except bouncy weapons, have an evolution.
  • Torrona's Box, added in patch 0.7.2, was not used for any evolution until the introduction of Sole Solution, added in patch 0.11.0.
    • It is also the only passive item that is involved with more than one evolution in the base v1.0 version of the game.
  • Bone is the first bouncy weapon to get an evolution, with an unique way to evolve by obtaining a relic, and leveling the original weapon owner to level 80.
  • SpellStrom is the first ever three weapon union.
  • Emegency Meeting is the first DLC that uses only DLC items to evolve weapons.
  • The passive items in Emergency Meeting resembles the character that uses the base weapon.

External links[]

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